Box-headed Blood-bee
Sphecodes monilicornis
This record describes Sphecodes monilicornis (species). Recorded family: Halictidae. Recorded order: Hymenoptera.
Identification and Appearance

One of three medium-sized to large Sphecodes (Blood-bees), in which females have punctures (rather than just rugosity) behind the ocelli. Males are one of a number of species with somewhat compressed and shiny antennal flagellar segments and are best confirmed using genitalia. S. monilicornis is easily separated from the other two (S. gibbus and S. reticulatus ) by the box-shaped head when seen in top view, with the hind corners well formed. The hairs of the hind tibiae are also whitish rather than dark like the other two, and the pygidium is broader. Most solitary bees and wasps are difficult to identify, and can rarely be identified from photos taken in the field. All red-rated records should include a photo or set of photos of the specimen, illustrating the key characters taken from a standard key, which should also be referenced (e.g. ‘Falk, 2015’). The full set of key characters are generally not visible in field photos and photos are rarely sharp enough. To aid in the verification of your records, please include face shot, side, top and wings. The notes should state whether male or female, and explain how the specimen met the key characters. Although NS may not be able to identify the species even if these reference photos are provided, the photos will be stored with the record and may allow it be identified in future. Alternatively, NS will accept records identified by a recognised local or national expert, or that have been identified via BWARS Facebook Group . If you have obtained this advice, please note the name of the person/organisation identifying the record in the ‘determiner’ field (e.g. ‘Stuart Roberts, BWARS Facebook’) rather than just a comment of ‘BWARS Facebook’. Females fly from late March to mid-September. Males from July to September. Cleptoparasite of several species of ground nesting bee including Lasioglossum calceatum, albipes and malachurum . Widespread and locally common in the southern half of Britain with records extending north to Scotland.
Classification
- Kingdom
- Animalia
- Phylum
- Arthropoda
- Class
- Insecta
- Order
- Hymenoptera
- Family
- Halictidae
- Genus
- Sphecodes
Imported taxonomy record. Source availability varies; see the source references and editorial policy below.
Habitat and Range
Habitat
Wherever the hosts are found.
Seasonal Activity
Females fly from late March to mid-September. Males from July to September.
Diet and Behavior
Life History
Cleptoparasite of several species of ground nesting bee including
Additional species information
Recorded status
Widespread and locally common in the southern half of Britain with records extending north to Scotland.
About Flies: the wider group
Order-level background for Hymenoptera, the group containing Sphecodes monilicornis. Species-specific details appear above.
Group overview
Hymenoptera includes over 150,000 described species of ants, bees, wasps, and sawflies. Earth harbors approximately 20 quadrillion (20×10^15) ants, with a combined biomass of ~12 megatons of dry carbon - exceeding the combined biomass of all wild birds and mammals (PNAS, 2022).
Ecological role of the group
About 35% of the world's food crops depend on animal pollinators. The chocolate industry, valued at over $100 billion, depends on pollinating flies and midges. Ants are the dominant soil engineers in most terrestrial ecosystems, turning more soil than earthworms in many habitats.
Economic and practical significance of the group
Pollinators contribute $235-577 billion worth of annual global food production (FAO). In the U.S., bee pollination alone benefits crops valued at $18 billion annually, and honey bees are responsible for $5.4 billion in agricultural productivity. Between April 2023-2024, 55.1% of managed honey bee colonies were lost. The U.S. bee population dropped from 5 million to 2.5 million.
Recorded sources for this group: FAO, USDA, EPA, PNAS (2022), US Fish & Wildlife Service. Imported estimates may describe particular regions or reporting periods.
Photos of Box-headed Blood-bee (8)
Frequently Asked Questions About Box-headed Blood-bee
What is Box-headed Blood-bee?
Box-headed Blood-bee (Sphecodes monilicornis) is a species in the family Halictidae, within the order Hymenoptera.
Where is Box-headed Blood-bee found?
Habitat: Wherever the hosts are found.
When is Box-headed Blood-bee active?
Box-headed Blood-bee: seasonal activity - Females fly from late March to mid-September. Males from July to September..
What is the life cycle of a Box-headed Blood-bee?
Cleptoparasite of several species of ground nesting bee including
FROM LOOKING IT UP TO GETTING TO KNOW IT
Make sense of the bugs at home.50 household bug profiles for you. Photos, coloring, and activities for curious kids.








